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The aquarium must be very large. Congratulations. The plates should fit together, but it's OK if they fit together only in pairs. The interface between two plates should have no bevel, thus allowing them to operate as one plate. You can hook the down tubes to a power head or just an air pump. It's possible to create too much turbulence, so the air pump may be better than the power heads. In either case, the down tubes will draw water from beneath the plates, which means it is filtered by the gravel substrate. That's a good thing, because the beneficial bacteria live in the substrate as well as plants, etc. A new aquarium setup needs to go through the nitrogen cycle; check our blogs for more details, and ask questions. We're here to help. Getting the setup right will avoid lots of problems in the future.